3.1 Classification Systems
Early botanists grouped plants using only a few visible traits. Over time three broad approaches β and several modern tools β developed.
| System | Basis | Proposed by / Note |
| Artificial | One or a few superficial vegetative characters (e.g. number of stamens) | Linnaeus; gave equal weight to vegetative & sexual traits, separated closely related groups |
| Natural | Natural affinities β many characters together, incl. internal features (anatomy, embryology, phytochemistry) | Bentham & Hooker |
| Phylogenetic | Evolutionary relationships β members of a group share a common ancestor | Modern systems; also use molecular data |
Newer tools
- Numerical Taxonomy β computer-based; each observable character is coded & numbered so hundreds of traits are weighed at once.
- Cytotaxonomy β based on cytological information: chromosome number, structure & behaviour.
- Chemotaxonomy β uses chemical constituents of the plant to resolve confusions.
Artificial (few traits)βNatural (many affinities)βPhylogenetic (evolution)
Big idea: Classification moved from "what looks similar" β to "what is truly related by evolution."
3.2 Algae
Algae are chlorophyll-bearing, simple, thalloid (no true rootβstemβleaf), largely aquatic autotrophs. The plant body is a thallus. They occur in fresh & marine water, moist stones, soil, wood, and even in/on other organisms.
Reproduction β three routes
- Vegetative β by fragmentation; each fragment grows into a new thallus.
- Asexual β mostly by zoospores (flagellated, motile) that germinate into new plants.
- Sexual β fusion of gametes:
| Type | Gametes | Example |
| Isogamous | Similar in size (both may be motile) | Ulothrix, Chlamydomonas (some) |
| Anisogamous | Dissimilar in size | Some species of Chlamydomonas |
| Oogamous | Large non-motile female + small motile male | Volvox, Fucus |
The three classes β compare carefully
| Feature | Chlorophyceae (Green) | Phaeophyceae (Brown) | Rhodophyceae (Red) |
| Major pigments | Chlorophyll a, b | Chl a, c + fucoxanthin | Chl a, d + phycoerythrin |
| Colour | Grass green | Olive-green β brown | Red |
| Stored food | Starch | Laminarin / Mannitol | Floridean starch |
| Cell wall | Cellulose | Cellulose + algin | Cellulose, pectin + polysulphate esters |
| Flagella | 2β8, equal, apical | 2, unequal, lateral | Absent |
| Habitat | Mostly freshwater | Mostly marine | Mostly marine (warmer waters) |
| Examples | Chlamydomonas, Volvox, Ulothrix, Spirogyra, Chara | Ectocarpus, Dictyota, Laminaria, Sargassum, Fucus | Polysiphonia, Porphyra, Gracilaria, Gelidium |
Economic & ecological importance
- Algae carry out at least half of Earth's COβ fixation (photosynthesis); they are primary producers of energy-rich food in aquatic food chains.
- Food: Porphyra, Laminaria, Sargassum.
- Hydrocolloids: algin (brown algae), carrageen (red algae). Agar from Gelidium & Gracilaria β culture media, jellies, ice-creams.
- Chlorella & Spirulina β protein-rich food supplements (even for space travellers).
Trap: Only red algae lack flagella completely. Green = starch; Brown = laminarin/mannitol; Red = floridean starch.
3.3 Bryophytes
Bryophytes are the "amphibians of the plant kingdom" β they live on land but need water to complete sexual reproduction. They lack true vascular tissue (no xylem/phloem).
- The main plant body is the gametophyte (haploid, dominant, photosynthetic), attached by rhizoids.
- Male organ = antheridium β flagellated antherozoids; female organ = archegonium (flask-shaped) β one egg.
- Water lets antherozoids swim to the archegonium β zygote.
- Zygote β sporophyte (diploid, partly dependent on gametophyte) β spores by meiosis β new gametophytes.
Spore (n)βGametophyte (n)βAntheridium + ArchegoniumβFertilisation (needs water)βZygote (2n)βSporophyte β meiosis β Spores
Liverworts
Moist, shady places; thalloid body (e.g. Marchantia). Asexual reproduction by gemmae (green multicellular buds in gemma cups) or fragmentation.
Mosses
Two stages: (1) protonema β creeping, green, branched filamentous stage from a spore; (2) leafy stage β upright, spirally arranged leaves, bears sex organs. Examples: Funaria, Polytrichum, Sphagnum.
Importance: Sphagnum (peat/bog moss) β peat (fuel) & water-holding packing for transport. Mosses form mats that reduce soil erosion and are pioneers on bare rock.
3.4 Pteridophytes
Pteridophytes are the first true land plants with vascular tissue (xylem & phloem). Examples: Selaginella, Equisetum, Pteris, Adiantum, Lycopodium, Salvinia, Marsilea.
- Dominant phase = sporophyte (2n) with true roots, stem & leaves.
- Leaves: small (microphylls, e.g. Selaginella) or large (macrophylls, ferns).
- Sporophytes bear sporangia on sporophylls; clusters of sporangia = sori.
- Sporangia β spores by meiosis β small, free-living, photosynthetic prothallus (gametophyte) needing cool, damp, shady soil.
- Gametophyte bears antheridia & archegonia; water needed for fertilisation.
Sporophyte (2n)βSporangia β meiosisβSpores (n)βProthallus (gametophyte)βFertilisation (water)βZygote β new Sporophyte
Homospory vs Heterospory
| Homosporous | Heterosporous |
| Spores | One kind of spore | Two kinds β large megaspores & small microspores |
| Examples | Most pteridophytes (Pteris, Dryopteris) | Selaginella, Salvinia |
| Significance | β | Megaspore β female gametophyte retained on parent β step toward the seed habit |
Heterospory is the precursor of the seed. Retention of the developing female gametophyte and zygote on the parent foreshadows seed plants.
3.5 Gymnosperms
Gymnosperms bear naked seeds β ovules are not enclosed in an ovary, so seeds are not covered by a fruit. Examples: Cycas, Pinus, Ginkgo, Cedrus, Ephedra.
- Small shrubs to giant trees (Sequoia, the redwood β among the tallest living organisms).
- Roots usually tap roots; Cycas has coralloid roots (Nβ-fixing cyanobacteria); Pinus has mycorrhiza.
- Leaves may be needle-like with thick cuticle & sunken stomata (xerophytic adaptations).
- Heterosporous β micro- & megaspores borne on cones / strobili.
- Male cone β microsporophylls + microsporangia β pollen grains (microspores).
- Female cone β megasporophylls + ovules; megaspore β female gametophyte with archegonia.
- No water needed β wind carries pollen to the ovule; pollen tube delivers male gametes.
- Zygote β embryo; ovule β naked seed (not inside a fruit).
Male cone β PollenβWind pollinationβOvule (female cone)βPollen tube β fertilisationβNaked seed
Key advance: The seed habit + wind pollination free gymnosperms from external water for reproduction.
3.6 Angiosperms
Angiosperms = flowering plants. Seeds develop inside an ovary that matures into a fruit. Sizes range from tiny Wolffia to giant Eucalyptus (~100 m).
- Stamen (male): anther's pollen sacs β pollen grains (microspores) β male gametes.
- Pistil/carpel (female): ovary β ovules; each ovule has an embryo sac (female gametophyte) with an egg + two polar nuclei.
- Double fertilisation (unique to angiosperms): male gamete + egg β zygote (2n) (syngamy); second male gamete + two polar nuclei β primary endosperm nucleus (3n) (triple fusion) β triploid endosperm.
Two classes
| Feature | Dicotyledons | Monocotyledons |
| Cotyledons in seed | Two | One |
| Leaf venation | Reticulate (net-like) | Parallel |
| Root system | Tap root | Fibrous root |
| Floral symmetry / parts | Usually in 4s or 5s (tetra/pentamerous) | In 3s (trimerous) |
| Examples | Pea, mustard, mango | Maize, wheat, grasses |
Syngamy: male gamete (n) + egg (n) β zygote (2n) | Triple fusion: male gamete (n) + 2 polar nuclei (2n) β endosperm (3n)
3.7 Life Cycles & Alternation of Generations
Plants alternate between a haploid gametophyte (makes gametes by mitosis) and a diploid sporophyte (makes spores by meiosis). Which phase dominates defines the life-cycle type.
| Type | Dominant phase | Other phase | Found in |
| Haplontic | Gametophyte (n), free-living | Only the zygote is diploid; zygote undergoes meiosis | Many algae β Volvox, Spirogyra, Chlamydomonas |
| Diplontic | Sporophyte (2n), free-living | Gametophyte reduced to a few cells; only gametes haploid | Seed plants (gymnosperms & angiosperms); alga Fucus |
| Haplo-diplontic | Both multicellular & free-living | One phase dominates the other | Bryophytes (gametophyte dom.) & pteridophytes (sporophyte dom.); Ectocarpus, Polysiphonia, Kelps |
Gametophyte (n) β gametes β fertilisation β Zygote (2n) β Sporophyte (2n) β meiosis β spores β Gametophyte (n) β¦
Memory anchor: Moss & ferns = "in-between" (haplo-diplontic). Bryophyte β gametophyte boss. Pteridophyte β sporophyte boss.